Listed from Konkian (lower Serravallian) deposits by Kolesnikov (1929). Later only mentioned from lower Sarmatian ... [details]
Volhynia, Podolia, Bessarabia, southern Ukraine, Crimea, Kerch Peninsula and central Ciscaucasia (no precise ... [details]
"Besskorbnaja" (Krasnodarskiy kray, Russia) (specified by Kolesnikov 1935) [details]
Uiratam, Mangghystau oblysy, Kazakhstan; Volhynian, early Sarmatian, Serravallian, middle Miocene [details]
Bessarabia (SW Ukraine or Moldova) [details]
Podolia, Bessarabia und southern Ukraine [details]
Bessarabia (SW Ukraine or Moldova) and Kerch Peninsula, Krym [details]
Bessarabia (SW Ukraine or Moldova), Kerch Peninsula (Krym) and central Ciscaucasia [details]