From editor or global species database
Type locality Shalfleet (Hamstead) ("Hempstead") and Totland (Headon Hill), Isle of Wight, and Milford-on-Sea (Hordwell Cliff) and Christchurch (Highcliffe) ("High Cliff"), Hampshire, United Kingdom; Priabonian to Lutetian to Rupelian, middle Eocene to early Oligocene [details]