original description
Locard, A. (1878). Description de la faune de la mollasse marine et d'eau douce du Lyonnais et du Dauphiné. <em>Archives du Musèum d'Histoire naturelle de Lyon.</em> 2, 1-284., available online at https://www.biodiversitylibrary.org/page/9660675
page(s): 216-217, pl. 19, figs 32-34 [details]
From editor or global species database
Type locality Hauterives, Drôme, France; Zanclean (early mammal zone MN 14), early Pliocene [details]