original description
Bogachev, V.V. (1961). Materialy k istorii presnovodnoi fauny Evrazii. <em>Izdatel'stvo Akademii Nauk Ukrainskoi SSR, Kiev.</em> 404 p.
page(s): 283, pl. 36, figs 9-14 [details] Available for editors [request]
From editor or global species database
Type locality Samara (Volch'ya Balka), Samarskaya oblast', Russian Federation; early Apsheronian, early Pleistocene [details]