original description
Grabau, A.W. (1923). Cretaceous fossils from Shantung. <em>Bulletin of the Geological Survey of China.</em> 5(2): 143-181.
page(s): 156-157, fig. 6a, b [details]
From editor or global species database
Type locality "in the gray shales in the lower part of the Wangshih series of Chiao-Hsien, east Shantung (loc. 220)" [Shandong Province, China] [details]