original description
Bogachev, V.V. (1961). Materialy k istorii presnovodnoi fauny Evrazii. <em>Izdatel'stvo Akademii Nauk Ukrainskoi SSR, Kiev.</em> 404 p.
page(s): 174-175, pl. 35, figs 1-7 [details] Available for editors
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From editor or global species database
Type locality "na vostochnom sklone Mugodzhar, pri vykhode rechki Chit-Irgiza iz polosy izverzhennykh porod na urochishche Kiziltashma" [on the eastern slope of the Mugodzhar hills, at the exit of the Shet-Yrghyz river from the strip of igneous rocks to the Kiziltashma tract] (Kazakhstan) [details]