original description
Yokoyama, M. (1922). Fossils from the Upper Musashino of Kazusa and Shimosa. <em>Journal of the College of Science, Imperial University of Tokyo.</em> 44(1): viii + 1-200, pls. 1-17 [7 July 1922]., available online at http://umdb.um.u-tokyo.ac.jp/DKoseibu/pdf/Ref_0465_.pdf
page(s): pp. 79-80; plate 4, fig. 4 [details]
From editor or global species database