original description
Oppenheim, P. (1919). Das Neogen in Kleinasien. <em>Zeitschrift der Deutschen Geologischen Gesellschaft.</em> 70, 1–210, 12 pls.
page(s): 134-135, pl. 10, fig. 1 [details]
From editor or global species database
Type locality Pamukkale (Giralan) ("Giralan bei Hierapolis"), Denizli, Turkey; lower Sazak Formation, Serravallian, middle Miocene [details]