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Bieler, R.; Petit, R. E. (2011). Catalogue of Recent and fossil “worm-snail” taxa of the families Vermetidae, Siliquariidae, and Turritellidae (Mollusca: Caenogastropoda). Zootaxa. 2948, 1-103.
163894
Bieler, R.; Petit, R. E.
2011
Catalogue of Recent and fossil “worm-snail” taxa of the families Vermetidae, Siliquariidae, and Turritellidae (Mollusca: Caenogastropoda)
Zootaxa
2948, 1-103
Publication
The taxonomy of the uncoiling “worm-snails” belonging to the marine gastropod families Vermetidae, Siliquariidae and Turritellidae is notoriously confused and their nominal species frequently mixed (in the literature as well as in type specimen collections) with members of superficially similar tube-building polychaete worms or members of unrelated molluscan groups. A long history of introducing and using infrasubspecific names and the rampant employment of homonymous names for unrelated taxa had contributed to a system that became unworkable. The current catalogue researches nearly 1,500 names that have been cited in conjunction with Recent and fossil taxa worm-snail taxonomy (six names above family-group level, 18 family-group names, 195 genus-group names, 1,278 species-group names). Each name’s validity and availability (in the sense of the I.C.Z.N. Code) was investigated and current placement within or outside the mentioned worm-snail families is suggested. 560 species-group names are interpreted as available for members of the worm-snail groups here under discussion. Of these, approximately 280 species-group names are available for extant taxa. Various formal First-Reviser actions are taken to resolve priority issues. The type species for Tulaxoda Blainville, 1828 is herein designated to be Serpulorbis polyphragma Sasso, 1827, making Tulaxoda an objective junior synonym of Thylacodes Guettard, 1770. Magilus Montfort, 1810 is declared a nomen protectum over Campulotus Guettard, 1770, a nomen oblitum. Recurring nomenclatural issues and those too complex to treat within the regular catalogue entries are discussed in 22 taxa notes. The catalogue is fully referenced in 766 literature titles and eight associated literature notes.
Biodiversity, Taxonomic and ecological diversity
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Aletes Carpenter, 1857 accepted as Thylacodes Guettard, 1770 (basis of record)
Aletes Carpenter, 1857 accepted as Thylacodes Guettard, 1770 (additional source)
Aletes centriquadrus var. imbricatus Carpenter, 1857 (additional source)
Aletes squamigerus Carpenter, 1857 accepted as Thylacodes squamigerus (Carpenter, 1857) (basis of record)
Angaria lacunosa Barnard, 1963 accepted as Stephopoma lacunosum (Barnard, 1963) (additional source)
Bivonia Gray, 1842 accepted as Dendropoma Mörch, 1861 (status source)
Bivonia albida Carpenter, 1875 (basis of record)
Bivonia compacta Carpenter, 1864 accepted as Petaloconchus compactus (Carpenter, 1864) (additional source)
Bivonia constrictor Mörch, 1862 accepted as Thylacodes constrictor (Mörch, 1862) (additional source)
Bivonia contorta Carpenter, 1857 accepted as Thylaeodus contortus (Carpenter, 1857) (basis of record)
Bivonia contorta var. indentata Carpenter, 1857 accepted as Thylaeodus indentatus (Carpenter, 1857) (basis of record)
Bivonia dragonella Okutani & Habe, 1975 accepted as Dendropoma dragonella (Okutani & Habe, 1975) (additional source)
Bivonia exserta Dall, 1881 (basis of record)
Bivonia petraea Monterosato, 1884 accepted as Dendropoma petraeum (Monterosato, 1884) accepted as Dendropoma cristatum (Biondi, 1859) (additional source)
Bivonia pteriae Parenzan, 1970 accepted as Vermetus granulatus (Gravenhorst, 1831) (basis of record)
Bivonia quoyi f. adspersa Mörch, 1862 (additional source)
Bivonia scopulorum Monterosato, 1917 accepted as Dendropoma scopulorum (Monterosato, 1917) (additional source)
Bivonia serpuloides Monterosato, 1892 (basis of record)
Bivonia subtriquetra Mörch, 1862 (basis of record)
Bivonia sutilis Mörch, 1862 accepted as Thylacodes sutilis (Mörch, 1862) (additional source)
Bivonia triquetra var. spirorbis Coen, 1937 (additional source)
Bivonia triquetra var. unicarinata Monterosato, 1917 accepted as Vermetus triquetrus Bivona-Bernardi, 1832 (additional source)
Campulotus Guettard, 1770 accepted as Magilus Montfort, 1810 (basis of record)
Caporbis africanus Bartsch, 1915 (basis of record)
Cerithiovermetus Bandel, 2006 (additional source)
Cerithiovermetus aqabensis Bandel, 2006 (additional source)
Cerithiovermetus vinxae Bandel, 2006 (source of synonymy)
Cladopoda Gray, 1850 accepted as Thylacodes Guettard, 1770 (additional source)
Cladopoda quoyi H. Adams & A. Adams, 1854 accepted as Vermetus quoyi (H. Adams & A. Adams, 1854) (additional source)
Dendropoma cristatum (Biondi, 1859) (additional source)
Dendropoma ghanaense Keen & J.E. Morton, 1960 (additional source)
Dendropoma gregaria [sic] accepted as Dendropoma gregarium Hadfield & Kay in Hadfield et al., 1972 (basis of record)
Dendropoma gregarium Hadfield & Kay in Hadfield et al., 1972 (additional source)
Dendropoma krypta [sic] accepted as Dendropoma kryptum S. M. Gardner, 1989 accepted as Cupolaconcha krypta (S. M. Gardner, 1989) (basis of record)
Dendropoma marchadi Keen & J.E. Morton, 1960 (additional source)
Dendropoma mejillonensis [sic] accepted as Dendropoma mejillonense Pacheco & Laudien, 2008 (additional source)
Dendropoma meroclista [sic] accepted as Dendropoma meroclistum Hadfield & Kay in Hadfield et al., 1972 accepted as Cupolaconcha meroclista (Hadfield & Kay in Hadfield et al., 1972) (additional source)
Dendropoma psarocephalum Hadfield & Kay in Hadfield et al., 1972 (additional source)
Dendropoma rhyssoconchum Hadfield & Kay in Hadfield et al., 1972 (additional source)
Dendropoma squamifera [sic] accepted as Dendropoma squamiferum Ponder, 1967 (basis of record)
Dendropoma tholia Keen & J.E. Morton, 1960 (additional source)
Dofania Mörch, 1860 (basis of record)
Eualetes Keen, 1971 (additional source)
Hummelinckiella Faber & Moolenbeek, 1999 (additional source)
Hummelinckiella borinquensis Faber & Moolenbeek, 1999 (additional source)
Lemintina Risso, 1826 (additional source)
Lemintina cuvieri Risso, 1826 (additional source)
Lemintina mcgintyi Olsson & Harbison, 1953 accepted as Petaloconchus mcgintyi (Olsson & Harbison, 1953) (additional source)
Lemintina pseudimbricata Nomura & Hatai, 1940 accepted as Dendropoma pseudimbricatum (Nomura & Hatai, 1940) (additional source)
Lemintina selecta var. arborea Monterosato, 1884 accepted as Thylacodes arenarius (Linnaeus, 1758) (additional source)
Lilax Finlay, 1926 accepted as Stephopoma Mörch, 1860 (additional source)
Macrophragma Carpenter, 1857 accepted as Petaloconchus H. C. Lea, 1843 (basis of record)
Magilina Vélain, 1877 (additional source)
Magilina serpuliformis Vélain, 1877 (additional source)
Novastoa Finlay, 1926 (additional source)
Petaloconchus H. C. Lea, 1843 (basis of record)
Petaloconchus apakadikike W. C. Kelly, 2007 (additional source)
Petaloconchus cereus Carpenter, 1857 (basis of record)
Petaloconchus cochlidium Carpenter, 1857 (basis of record)
Petaloconchus complicatus Dall, 1908 (additional source)
Petaloconchus flavescens Carpenter, 1857 (additional source)
Petaloconchus floridana [sic] accepted as Petaloconchus floridanus Olsson & Harbison, 1953 (basis of record)
Petaloconchus innumerabilis Pilsbry & Olsson, 1935 (additional source)
Petaloconchus interliratus Stearns, 1893 (additional source)
Petaloconchus keenae Hadfield & Kay, 1972 (additional source)
Petaloconchus lamellosus Ladd, 1972 (additional source)
Petaloconchus lilandikike W. C. Kelly, 2007 (additional source)
Petaloconchus macrophragma Carpenter, 1857 (additional source)
Petaloconchus merkana Ladd, 1972 (additional source)
Petaloconchus montereyensis Dall, 1919 (additional source)
Petaloconchus myrakeenae Absalão & Rios, 1987 (additional source)
Petaloconchus nerinaeoides Carpenter, 1857 (basis of record)
Petaloconchus octosectus Carpenter, 1857 (basis of record)
Petaloconchus renisectus Carpenter, 1857 (basis of record)
Petaloconchus renisectus var. gordialis Mörch, 1862 accepted as Petaloconchus renisectus Carpenter, 1857 (additional source)
Petaloconchus renisectus var. woodwardii Carpenter, 1857 (basis of record)
Petaloconchus sculpturatus H. C. Lea, 1843 † (basis of record)
Petaloconchus tokyoensis (Pilsbry, 1895) (additional source)
Petalopoma Schiaparelli, 2002 (additional source)
Serpula conica Dillwyn, 1817 accepted as Vermetus conicus (Dillwyn, 1817) (additional source)
Serpula intestinalis Gmelin, 1791 accepted as Vermetus intestinalis (Gmelin, 1791) (additional source)
Serpula lumbricalis Linnaeus, 1758 accepted as Vermicularia lumbricalis (Linnaeus, 1758) (status source)
Serpula panamensis Chenu, 1846 accepted as Vermetidae incertae sedis panamensis (Chenu, 1846) (additional source)
Serpula tortuosa [Lightfoot], 1786 accepted as Vermicularia tortuosa (Lightfoot, 1786) (additional source)
Serpula turboides Chiereghini in Nardo, 1847 accepted as Siphonium gaederopi Mörch, 1861 accepted as Dendropoma gaederopi (Mörch, 1861) (additional source)
Serpulorbis Sasso, 1827 accepted as Thylacodes Guettard, 1770 (additional source)
Serpulorbis aotearoicus J. E. Morton, 1951 accepted as Thylacodes aotearoicus (J. E. Morton, 1951) (additional source)
Serpulorbis aureus Hughes, 1978 accepted as Thylacodes aureus (Hughes, 1978) (additional source)
Serpulorbis birugosus Weisbord, 1962 accepted as Hydroides bispinosus Bush, 1910 accepted as Hydroides bispinosa Bush, 1910 [original] (basis of record)
Serpulorbis catella Weisbord, 1962 accepted as Pomatoceros minutus Rioja, 1941 accepted as Spirobranchus minutus (Rioja, 1941) (additional source)
Serpulorbis catellus Weisbord, 1962 accepted as Pomatoceros minutus Rioja, 1941 accepted as Spirobranchus minutus (Rioja, 1941) (additional source)
Serpulorbis daidai Scheuwimmer & Nishiwaki, 1982 accepted as Thylacodes daidai (Scheuwimmer & Nishiwaki, 1982) (additional source)
Serpulorbis hadfieldi W. C. Kelly, 2007 accepted as Thylacodes hadfieldi (W. C. Kelly, 2007) (additional source)
Serpulorbis lamarckii Vaillant, 1871 accepted as Thylacodes lamarckii (Vaillant, 1871) (additional source)
Serpulorbis polyphragma Sasso, 1827 accepted as Thylacodes arenarius (Linnaeus, 1758) (basis of record)
Serpulorbis squamolineatus Petuch, 2002 accepted as Thylacodes squamolineatus (Petuch, 2002) (additional source)
Serpulorbis trimeresurus Shikama & Horikoshi, 1963 accepted as Thylacodes trimeresurus (Shikama & Horikoshi, 1963) (status source)
Serpulorbis validus Kuroda & Habe, 1952 (status source)
Serpulorbis variabilis Hadfield & Kay, 1972 accepted as Thylacodes variabilis (Hadfield & Kay, 1972) (additional source)
Serpulorbis varidus Okutani & Habe, 1975 accepted as Thylacodes varidus (Okutani & Habe, 1975) (status source)
English worm-snails for Vermetidae Rafinesque, 1815
 Authority

Attributed to Oyama in the original, based on manuscript data. [details]

 Authority

Erroneously credited to other authors. See discussion in Bieler & Petit (2011: 58 and taxa note 21 in that work). [details]

 Homonymy

A junior primary homonym of Vermetus polygonus corrugatus Cossmann in Cossmann & Pissaro, 1910 (Eocene). ... [details]

 Homonymy

Not to be confused with junior primary homonyms Vermetus cristatus Sandberger, 1860 and V. cristatus K. Martin, ... [details]

 Homonymy

A junior primary homonym of V. imbricatus Sandberger, 1859 and of V. imbricatus Dunker,1860; permanently invalid under ... [details]

 Homonymy

See Vermetus imbricatus Sandberger, 1859 (Oligocene). [details]

 Homonymy

A junior primary homonym of Vermetus indicus Daudin, 1800. Name may be preserved under ICZN (1999) Art. 23.9.5 if ... [details]

 Homonymy

In need of investigation: A junior secondary homonym of Vermicularia costalis (Lamarck, 1818).  [details]

 Identification

Turritellidae? [details]

 Nomenclature

Genus name placed on the Official List in Opinion 1425 (I.C.Z.N. 1987) [details]

 Nomenclature

Placed on the Official List of Specific Names in Zoology by Opinion 1425 (I.C.Z.N. 1987). [details]

 Nomenclature

Originally spelled 5-costatus. [details]

 Spelling

Emendation from the male ending -i by Bieler & Petit (2011: 67) [details]

 Spelling

Name ending corrected to reflect male gender of generic name. [details]

 Spelling

Genus misspelled "Serpuorbis" in original. [details]

 Spelling

Originally introduced as Strephopoma [sic] nucleocostata [details]

 Spelling

As "balani-tintinnabuli".  [details]

 Spelling

Originally spelled "retifera". [details]

 Spelling

Originally spelled "albina", as a variety in Vermetus granuatus [sic].  [details]

 Spelling

Originally spelled "picta", as a variety in Vermetus granuatus [sic].  [details]

 Spelling

Originally spelled "albina". [details]

 Spelling

Bivona-Bernardi introduced the name as Vermetus triquetrus, using the correctly formed Latin suffix –quetrus, ... [details]

 Status

Established by Carpenter in synonymy, which would make it unavailable under I.C.Z.N. (1999) Art. 11.6. However, its ... [details]

 Status

Introduced as "nom. prov." although with a description. It was recognized by Keen (1961: 198) as Petaloconchus ... [details]

 Status

Kuroda & Habe (1952) concluded that Pilsbry had figured two species as Thylacodes medusae and introduced the new ... [details]

 Status

Temporarily used to accommodate taxa that are deemed to be siliquariids but have not been assigned to a genus. ... [details]

 Status

Type not designated. Generic name is invalid because preoccupied by Siphonium Link, 1807 [Cephalopoda]. For a ... [details]

 Status

Temporarily used to accommodate taxa that are deemed to be vermetids but have not been assigned to a genus. These ... [details]

 Status

Described in polychaete worm genus; current genus-level placement in Vermetidae uncertain. [details]

 Status

Described in Vermicularia (Turritellidae); current genus-level placement in Vermetidae uncertain. [details]

 Status

Described in polychaete worm genus; current genus-level placement in Vermetidae uncertain. [details]

 Status

Described in polychaete worm genus; current genus-level placement in Vermetidae uncertain. [details]

 Status

Described in polychaete worm genus; current genus-level placement in Vermetidae uncertain. [details]

 Status

Described in polychaete worm genus; current genus-level placement in Vermetidae uncertain. [details]

 Status

Described in polychaete worm genus; current genus-level placement in Vermetidae uncertain. [details]

 Status

A junior primary homonym of Vermetus semisurrectus var. minor Monterosato, 1892; permanently invalid under ICZN (1999) ... [details]

 Status

Described in polychaete worm genus; current genus-level placement in Vermetidae uncertain. [details]

 Status

Originally infrasubspecific but treated as subspecies by Turton (1932: 127). Bieler (1996: 25) credited Turton as ... [details]

 Status

A junior primary homonym of Vermetus semisurrectus var. minor Monterosato, 1892; permanently invalid under ICZN (1999) ... [details]

 Status

A junior primary homonym of Vermetus semisurrectus var. minor Monterosato, 1892; permanently invalid under ICZN (1999) ... [details]

 Status

A junior primary homonym of Vermetus semisurrectus var. minor Monterosato, 1892; permanently invalid under ICZN (1999) ... [details]

 Status

Treated as polychaete by Bieler & Petit (2011: 41). [details]

 Status

Mörch (1871: 131) stated that it might not be based on a mollusk. Placed in synonymy of Vermetus (Bivonia) ... [details]

 Status

Treated as a polychaete by Bieler & Petit (2011: 53). [details]

 Status

Described in polychaete worm genus; current genus-level placement in Vermetidae uncertain. [details]

 Status

Described in polychaete worm genus; current genus-level placement in Vermetidae uncertain. [details]

 Status

Boettger mistakenly renamed the senior homonym, Vermetus imbricatus Sandberger, "1863" [error for 1859], non V. ... [details]

 Status

Described in Vermicularia(/i> (Turritellidae); current genus-level placement in Vermetidae uncertain. [details]

 Status

Originally spelled "bicarinata", taking the gender of "forma". A junior primary homonym of Vermetus bicarinatus ... [details]

 Status

Junior primary homonym of Vermetus granulatus forma repens in the same work by Monterosato. [details]

 Synonymy

Keen (1961: 198) considered Magilina a possible synonym of Dendropoma[details]

 Synonymy

A justified emendation of Tulaxodus Guettard, 1770 [details]

 Taxonomic remark

Described in polychaete worm genus. [details]

 Taxonomy

Ex Kuroda manuscript. Bivonia dragonella was first described by Kuroda in 1947 in the Yume-hamaguri, a journal ... [details]

 Taxonomy

Stated by Kuroda & Habe (1952: 62) to be "a tube of an Annelid" but placed in Dendropoma by Habe & Masuda (1990: 23). [details]

 Taxonomy

Originally infrasubspecific but treated as subspecies by Iwakawa (1909: 79). Bieler (1996: 25) credited Iwakawa as ... [details]

 Taxonomy

Kuroda & Habe (1952) concluded that Pilsbry had figured two species as Thylacodes medusae and introduced the new ... [details]

 Taxonomy

Originally spelled "angulata". A junior primary homonym of Vermetus angulatus Rousseau in Chenu, 1844. See Bieler & ... [details]

 Taxonomy

New name introduced for Vermetus sipho Rousseau in Chenu, 1844, non Lamarck, 1818 [details]

 Taxonomy

Unnecessary new name for Vermicularia nodosa Hedley, 1907, non Vermetus nodosus Kaunhowen, 1897 [sic; = 1898]. [details]

 Taxonomy

Incorrectly considered to be a junior homonym of Vermetus nodosa Kaunhowen, 1898 and renamed Vermicularia hedleyi ... [details]

 Type designation

Type species, by subsequent designation of Kobelt (1878: 139), Vermetus arenarius ‘Quoy (nec. L.)’ which was ... [details]


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