original description
Babak, E.V. (1983). Pliotsenovyye i chetvertichnyye dreyssenidy Evskinskogo basseyna. Trudy Paleontologicheskogo Instituta, 204: 1-104, available online at http://www.geokniga.org/books/14107
page(s): 45-46, pl. 6, figs 12-16 [details]
From editor or global species database
Type locality "Zapadnaya Gruziya, r. Duab" [western Georgia, river Duab]; middle-late Kimmerian, Pliocene [details]