original description
Olsson A.A. (1931). Contribution to the Tertiary paleontology of northern Peru: part 4, the Peruvian Oligocene. <em>Bulletins of American Paleontology.</em> 17(63): 97-264 [1-168], pls 13-33., available online at https://www.biodiversitylibrary.org/page/32443892
page(s): 182, pl. 15, figs 9, 11 [details]
From editor or global species database
Type locality Chira valley, Lagunitas, Caleto Sal, Punta Mero (Peru) and Que. Charura (northern Colombia) [details]