original description
Nagao, T. (1928). Palaeogene fossils of the Island of Kyûshû, Japan, Part II. <em>Science Reports of the Tôhoku Imperial University, 2nd Series, Geology.</em> 12(1): 11-140, pl. 1-17.
page(s): 103, pl. 15, fig. 2-2b [details] Available for editors [request]
From editor or global species database
Type locality The Kamiyamada fossil bed; the shaft of the Kamiyamada Mine, Yamada-machi, Kaho-gun, province of Chikuzen (Kyushu, Japan) [details]