original description
Seninski, K. (1905). Novyya Dannyya o neogenovych' plastach' Yugozapadnogo Zakavkaz'ya. <em>Trudy Obshchestva Estestvoispytateley pri Imperatorskom Yuryevskom Universitete.</em> 16, 1-80.
page(s): 50, pl. 2, figs 19-20 [details]
source of synonymy
Taktakishvili, I.G. (1973). Pliotsenovyye dreyssenidy Zapadnoy Gruzii. Metsniyereba, Tbilisi, 150 pp., available online at https://www.geo-fund.am/files/library/1/15229236051436.pdf
page(s): 71 [details]
source of synonymy
Babak, E.V. (1983). Pliotsenovyye i chetvertichnyye dreyssenidy Evskinskogo basseyna. Trudy Paleontologicheskogo Instituta, 204: 1-104, available online at http://www.geokniga.org/books/14107
page(s): 39 [details]
From editor or global species database
Type locality "Reka Duab" [River Duab] (Abkhazia) [details]