original description
Seninski, K. (1905). Novyya Dannyya o neogenovych' plastach' Yugozapadnogo Zakavkaz'ya. <em>Trudy Obshchestva Estestvoispytateley pri Imperatorskom Yuryevskom Universitete.</em> 16, 1-80.
page(s): 43, pl. 2, figs 23-24 [details]
From editor or global species database
Type locality "Reka Duab" [River Duab] (Abkhazia) [details]