MolluscaBase taxon details
Pupilla J. Fleming, 1828
426395 (urn:lsid:marinespecies.org:taxname:426395)
accepted
Genus
Pupa marginata Draparnaud, 1801 accepted as Pupilla muscorum (Linnaeus, 1758) (type by monotypy)
Clausilia (Pupa) Draparnaud, 1801 · unaccepted > junior subjective synonym
Glischrus (Pupa) Draparnaud, 1801 · unaccepted
Paracoryna Flach, 1890 † · unaccepted > junior subjective synonym
Pupa Draparnaud, 1801 · unaccepted (invalid: junior homonym of Pupa...)
invalid: junior homonym of Pupa Röding, 1798
Pupa (Paracoryna) Flach, 1890 † · unaccepted > junior subjective synonym
Pupa (Pupilla) J. Fleming, 1828 · unaccepted
Pupilla (Afripupilla) Pilsbry, 1921 · alternative representation
Pupilla (Fragipupilla) Schileyko, 1984 · alternative representation
Pupilla (Pupilla) J. Fleming, 1828 · alternative representation
Pupilla (Striopupilla) Pilsbry, 1921 · alternative representation
Sphyradium (Pupa) Draparnaud, 1801 · unaccepted > superseded combination
Torquatella Held, 1838 · unaccepted (junior synonym)
Vertigo (Pupa) Draparnaud, 1801 · unaccepted (unaccepted combination)
- Species Pupilla aeoli (Hilber, 1884) †
- Species Pupilla afghanicum Solem, 1979
- Species Pupilla alabiella Schileyko, 1984
- Species Pupilla alaskensis Nekola & Coles, 2014
- Species Pupilla alluvionica Meng & L. Hoffman, 2008
- Species Pupilla almuhambetovae Prysjazhnjuk, 2017 †
- Species Pupilla alpicola (Charpentier, 1837)
- Species Pupilla altaica Meng & L. Hoffman, 2008
- Species Pupilla annandalei Pilsbry, 1921
- Species Pupilla anzobica Izzatullaev, 1970
- Species Pupilla armeniaca (Issel, 1865)
- Species Pupilla bargi Prysiazhniuk, 2017 †
- Species Pupilla belokrysi Steklov, 1966 †
- Species Pupilla bipapulata Akramowski, 1943
- Species Pupilla bituberculata H.-J. Wang & S.-Y. Guo, 1991 †
- Species Pupilla blainvilleana (Dupuy, 1850) †
- Species Pupilla blandii E. S. Morse, 1865
- Species Pupilla bogdanovkaense Prysiazhniuk, 2017 †
- Species Pupilla capitani (de Morgan, 1920) †
- Species Pupilla crossei (Michaud, 1862) †
- Species Pupilla cupella O. Boettger, 1889 †
- Species Pupilla diezi (Flach, 1890) †
- Species Pupilla duplicata (Preston, 1911)
- Species Pupilla eurina (W. H. Benson, 1864)
- Species Pupilla ficulnea (Tate, 1894)
- Species Pupilla gallae Tzvetkov, 1940
- Species Pupilla goniodon Pilsbry, 1927
- Species Pupilla grabaui (Ping, 1929) †
- Species Pupilla guadalupensis Pilsbry, 1927
- Species Pupilla gutta (W. H. Benson, 1864)
- Species Pupilla hebes (Ancey, 1881)
- Species Pupilla hokkaidoensis Nekola, Coles & S. Chiba, 2014
- Species Pupilla hopeiensis (Ping, 1929) †
- Species Pupilla hudsoniana Nekola & Coles, 2014
- Species Pupilla iheringi Suter, 1900
- Species Pupilla impressa (F. Sandberger, 1862) †
- Species Pupilla inequidenta Schileyko & Almuhambetova, 1979
- Species Pupilla inermis L. S. Russell, 1931 †
- Species Pupilla inops (Reinhardt, 1877)
- Species Pupilla iratiana (Dupuy, 1850) †
- Species Pupilla khunjerabica Auffenberg & Pokryszko, 2009
- Species Pupilla kyrostriata Walther & Hausdorf, 2014
- Species Pupilla limata Schileyko, 1984
- Species Pupilla loessica Ložek, 1954
- Species Pupilla michaudi Wenz, 1919 †
- Species Pupilla mlomnickii (Friedberg, 1905) †
- Species Pupilla muscorum (Linnaeus, 1758)
- Species Pupilla mutabilis Steklov, 1966 †
- Species Pupilla paraturcmenica Hlaváč & Pokryszko, 2009
- Species Pupilla parvula (Deshayes, 1863) †
- Species Pupilla perlabiata Gottschick & Wenz, 1919 †
- Species Pupilla poltavica O. Boettger, 1889 †
- Species Pupilla profundotuberculata H.-J. Wang & S.-Y. Guo, 1991 †
- Species Pupilla quxuensis D.-N. Chen & G.-Q. Zhang, 2001
- Species Pupilla retusa (F. Sandberger, 1858) †
- Species Pupilla roberti Prysiazhniuk, 2017 †
- Species Pupilla salemensis (W. T. Blanford & H. F. Blanford, 1861)
- Species Pupilla satparanica Pokryszko & Auffenberg, 2009
- Species Pupilla selecta (Thomä, 1845) †
- Species Pupilla selectiformis (Simionescu & Barbu, 1940) †
- Species Pupilla shantungensis Yen, 1969 †
- Species Pupilla signataeformis Steklov, 1966 †
- Species Pupilla simplexa Y.-T. Li, 1985 †
- Species Pupilla sonorana (Sterki, 1899)
- Species Pupilla staszicii (Łomnicki, 1886) †
- Species Pupilla steinheimensis (K. Miller, 1900) †
- Species Pupilla sterkiana (Pilsbry, 1889)
- Species Pupilla sterrii (Voith, 1840)
- Species Pupilla striopolita Schileyko, 1984
- Species Pupilla subquadridens (A. d'Orbigny, 1852) †
- Species Pupilla syngenes (Pilsbry, 1890)
- Species Pupilla talassika Meng, 2008
- Species Pupilla tetrodus (O. Boettger, 1870)
- Species Pupilla triplicata (S. Studer, 1820)
- Species Pupilla triplicatoidea Steklov, 1966 †
- Species Pupilla tschumakovi Prysjazhnjuk, 2017 †
- Species Pupilla turcmenia O. Boettger, 1889
- Species Pupilla zeisslerae Schlickum, 1975 †
- Species Pupilla ziaratana Pokryszko & Auffenberg, 2009
- Subgenus Pupilla (Gastrocopta) Wollaston, 1878 accepted as Gastrocopta Wollaston, 1878 (unaccepted rank)
- Species Pupilla (Gastrocopta) munita (Reibisch, 1893) accepted as Gastrocopta munita (Reibisch, 1893) (unaccepted > superseded combination)
- Species Pupilla (Gastrocopta) reibischi Dall & Ochsner, 1928 accepted as Gastrocopta reibischi (Dall & Ochsner, 1928) (unaccepted > superseded combination)
- Subgenus Pupilla (Gibbulinopsis) Germain, 1919 accepted as Gibbulinopsis Germain, 1919 (considered as a separate genus)
- Species Pupilla (Gibbulinopsis) cryptodon (Heude, 1880) accepted as Gibbulinopsis cryptodon (Heude, 1880)
- Species Pupilla (Gibbulinopsis) fontana (Krauss, 1848) accepted as Gibbulinopsis fontana (Krauss, 1848) (unaccepted > superseded combination)
- Species Pupilla (Gibbulinopsis) signata (Mousson, 1873) accepted as Gibbulinopsis signata (Mousson, 1873) (unaccepted > superseded combination)
- Species Pupilla (Gibbulinopsis) blainvilleana (Dupuy, 1850) † represented as Pupilla blainvilleana (Dupuy, 1850) †
- Subgenus Pupilla (Omegapilla) Iredale, 1937 accepted as Omegapilla Iredale, 1937
- Species Pupilla (Omegapilla) australis (A. Adams & Angas, 1864) accepted as Omegapilla australis (A. Adams & Angas, 1864) (unaccepted > superseded combination)
- Subgenus Pupilla (Primipupilla) Pilsbry, 1921 accepted as Gibbulinopsis (Primipupilla) Pilsbry, 1921 represented as Gibbulinopsis Germain, 1919
- Species Pupilla (Primipupilla) selecta (Thomä, 1845) † accepted as Pupilla selecta (Thomä, 1845) †
- Species Pupilla (Primipupilla) signata (Mousson, 1873) accepted as Gibbulinopsis signata (Mousson, 1873) (unaccepted > superseded combination)
- Species Pupilla (Primipupilla) staszici [sic] † accepted as Pupilla staszicii (Łomnicki, 1886) † (unaccepted > misspelling - incorrect subsequent spelling)
- Species Pupilla barrackporensis Gude, 1914 accepted as Insulipupa brevicostis (W. H. Benson, 1849) (unaccepted > junior subjective synonym)
- Species Pupilla blandi [sic] accepted as Pupilla blandii E. S. Morse, 1865 (unaccepted > misspelling - incorrect subsequent spelling)
- Species Pupilla brevicostis (W. H. Benson, 1849) accepted as Insulipupa brevicostis (W. H. Benson, 1849) (unaccepted > superseded combination)
- Species Pupilla cinghalensis Gude, 1914 accepted as Indopupa cinghalensis (Gude, 1914) (original combination)
- Species Pupilla cryptodon (Heude, 1880) accepted as Gibbulinopsis cryptodon (Heude, 1880) (unaccepted combination)
- Species Pupilla cupa (Jan, 1832) accepted as Pupilla muscorum (Linnaeus, 1758) (unaccepted > junior subjective synonym)
- Species Pupilla diopsis (W. H. Benson, 1863) accepted as Vertigo diopsis (W. H. Benson, 1863) (unaccepted > superseded combination)
- Species Pupilla draparnaudii Leach, 1852 accepted as Lauria cylindracea (Da Costa, 1778) (unaccepted > junior subjective synonym)
- Species Pupilla floridana Dall, 1885 accepted as Gastrocopta pentodon (Say, 1822) (unaccepted > junior subjective synonym)
- Species Pupilla fontana (Krauss, 1848) accepted as Gibbulinopsis fontana (Krauss, 1848) (unaccepted > superseded combination)
- Species Pupilla gracilis Izzatullaev, 1970 accepted as Gibbulinopsis gracilis (Izzatullaev, 1970) (original combination)
- Species Pupilla heudeana (Möllendorff, 1884) accepted as Gibbulinopsis cryptodon cryptodon (Heude, 1880) (unaccepted > junior subjective synonym)
- Species Pupilla hudsonianum [sic] accepted as Pupilla hudsoniana Nekola & Coles, 2014 (wrong gender agreement of specific epithet)
- Species Pupilla muscerda (W. H. Benson, 1853) accepted as Microstele muscerda (W. H. Benson, 1853) (unaccepted combination)
- Species Pupilla paulinoi Locard, 1899 accepted as Leiostyla anglica (A. Férussac, 1821) (unaccepted > junior subjective synonym)
- Species Pupilla praesignata Steklov † (unaccepted > nomen nudum)
- Species Pupilla pratensis (Clessin, 1871) accepted as Pupilla alpicola (Charpentier, 1837) (unaccepted > junior subjective synonym)
- Species Pupilla pseudotriplicata Steklov † (unaccepted > nomen nudum)
- Species Pupilla quadrigranata (F. Sandberger, 1858) † accepted as Pupilla selecta (Thomä, 1845) † (unaccepted > junior subjective synonym)
- Species Pupilla reibischi Dall & Ochsner, 1928 accepted as Gastrocopta reibischi (Dall & Ochsner, 1928) (original name)
- Species Pupilla salmensis (W. T. Blanford & H. F. Blanford, 1861) accepted as Pupilla salemensis (W. T. Blanford & H. F. Blanford, 1861) (misspelling of original name.)
- Species Pupilla seriola (W. H. Benson, 1863) accepted as Vertigo seriola (W. H. Benson, 1863) (unaccepted > superseded combination)
- Species Pupilla signata (Mousson, 1876) accepted as Gibbulinopsis signata (Mousson, 1873) (unaccepted > superseded combination)
- Species Pupilla stoneri Chamberlin & D. T. Jones, 1929 accepted as Gastrocopta pilsbryana (Sterki, 1890) (unaccepted > junior subjective synonym)
- Species Pupilla submuscorum Gottschick & Wenz, 1919 † accepted as Gibbulinopsis submuscorum (Gottschick & Wenz, 1919) † (unaccepted > superseded combination)
- Species Pupilla turcmenica [sic!] accepted as Pupilla turcmenia O. Boettger, 1889 (unaccepted > misspelling - incorrect original spelling)
- Species Pupilla wolfae Tzvetkov, 1940 accepted as Pupilla sterrii (Voith, 1840) (junior synonym)
- Subgenus Pupilla (Afripupilla) Pilsbry, 1921 represented as Pupilla J. Fleming, 1828
- Species Pupilla (Afripupilla) tetrodus (O. Boettger, 1870) represented as Pupilla tetrodus (O. Boettger, 1870)
- Subgenus Pupilla (Fragipupilla) Schileyko, 1984 represented as Pupilla J. Fleming, 1828
- Species Pupilla (Fragipupilla) alabiella Schileyko, 1984 represented as Pupilla alabiella Schileyko, 1984 (basionym)
- Species Pupilla (Fragipupilla) limata Schileyko, 1984 represented as Pupilla limata Schileyko, 1984 (basionym)
- Subgenus Pupilla (Pupilla) J. Fleming, 1828 represented as Pupilla J. Fleming, 1828
- Species Pupilla (Pupilla) anodon (Deshayes, 1863) † accepted as Negulopsis anodon (Deshayes, 1863) † (unaccepted > superseded combination, tentative superseded combination)
- Species Pupilla (Pupilla) pratensis (Clessin, 1871) accepted as Pupilla (Pupilla) alpicola (Charpentier, 1837) represented as Pupilla alpicola (Charpentier, 1837) (unaccepted > junior subjective synonym)
- Species Pupilla (Pupilla) striopolita Schileyko, 1984 accepted as Pupilla striopolita Schileyko, 1984
- Species Pupilla (Pupilla) alpicola (Charpentier, 1837) represented as Pupilla alpicola (Charpentier, 1837)
- Species Pupilla (Pupilla) annandalei Pilsbry, 1921 represented as Pupilla annandalei Pilsbry, 1921
- Species Pupilla (Pupilla) bigranata (Rossmässler, 1839) represented as Pupilla bigranata (Rossmässler, 1839)
- Species Pupilla (Pupilla) cupella O. Boettger, 1889 † represented as Pupilla cupella O. Boettger, 1889 †
- Species Pupilla (Pupilla) eurina (W. H. Benson, 1864) represented as Pupilla eurina (W. H. Benson, 1864)
- Species Pupilla (Pupilla) hebes (Ancey, 1881) represented as Pupilla hebes (Ancey, 1881)
- Species Pupilla (Pupilla) inops (Reinhardt, 1877) represented as Pupilla inops (Reinhardt, 1877)
- Species Pupilla (Pupilla) khunjerabica Auffenberg & Pokryszko, 2009 represented as Pupilla khunjerabica Auffenberg & Pokryszko, 2009
- Species Pupilla (Pupilla) muscorum (Linnaeus, 1758) represented as Pupilla muscorum (Linnaeus, 1758)
- Species Pupilla (Pupilla) paraturcmenica Hlaváč & Pokryszko, 2009 represented as Pupilla paraturcmenica Hlaváč & Pokryszko, 2009
- Species Pupilla (Pupilla) satparanica Pokryszko & Auffenberg, 2009 represented as Pupilla satparanica Pokryszko & Auffenberg, 2009
- Species Pupilla (Pupilla) sterrii (Voith, 1840) represented as Pupilla sterrii (Voith, 1840)
- Species Pupilla (Pupilla) triplicata (S. Studer, 1820) represented as Pupilla triplicata (S. Studer, 1820)
- Species Pupilla (Pupilla) turcmenica O. Boettger, 1889 represented as Pupilla turcmenia O. Boettger, 1889
- Species Pupilla (Pupilla) zeisslerae Schlickum, 1975 † represented as Pupilla zeisslerae Schlickum, 1975 †
- Species Pupilla (Pupilla) ziaratana Pokryszko & Auffenberg, 2009 represented as Pupilla ziaratana Pokryszko & Auffenberg, 2009
- Subgenus Pupilla (Striopupilla) Pilsbry, 1921 represented as Pupilla J. Fleming, 1828
- Species Pupilla (Striopupilla) goniodon Pilsbry, 1927 represented as Pupilla goniodon Pilsbry, 1927
- Species Pupilla (Striopupilla) guadalupensis Pilsbry, 1927 represented as Pupilla guadalupensis Pilsbry, 1927
- Species Pupilla (Striopupilla) sterkiana (Pilsbry, 1889) represented as Pupilla sterkiana (Pilsbry, 1889)
- Species Pupilla bigranata (Rossmässler, 1839) (uncertain > taxon inquirendum)
- Species Pupilla bogdanovi Urbanski, 1960 (uncertain)
- Species Pupilla chinensis (Hilber, 1884) † (uncertain > taxon inquirendum)
- Species Pupilla elegans (Matheron, 1832) † (uncertain > taxon inquirendum)
- Species Pupilla l'eprevieri Pallary, 1928 (uncertain > taxon inquirendum)
- Species Pupilla richthofeni (Hilber, 1884) † (uncertain > taxon inquirendum)
- Species Pupilla scapula (F. Sandberger, 1875) † (uncertain > taxon inquirendum)
- Species Pupilla valkanovi Urbanski, 1960 (uncertain)
feminine
Fleming, J. (1828). A history of British animals, exhibiting the descriptive characters and systematical arrangement of the genera and species of quadrupeds, birds, reptiles, fishes, Mollusca, and Radiata of the United Kingdom; including the indigenous, extirpated, and extinct kinds, together with periodical and occasional visitants. <em>Edinburgh, Bell & Bradfute / London, James Duncan.</em> Pp. i-xxii, 1-565. corrigenda., available online at http://www.biodiversitylibrary.org/item/18569
page(s): 268; note: name attributed by Fleming to Leach (MS). [details]
page(s): 268; note: name attributed by Fleming to Leach (MS). [details]
MolluscaBase eds. (2025). MolluscaBase. Pupilla J. Fleming, 1828. Accessed at: https://www.molluscabase.org/aphia.php?p=taxdetails&id=426395 on 2025-06-30
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original description
Fleming, J. (1828). A history of British animals, exhibiting the descriptive characters and systematical arrangement of the genera and species of quadrupeds, birds, reptiles, fishes, Mollusca, and Radiata of the United Kingdom; including the indigenous, extirpated, and extinct kinds, together with periodical and occasional visitants. <em>Edinburgh, Bell & Bradfute / London, James Duncan.</em> Pp. i-xxii, 1-565. corrigenda., available online at http://www.biodiversitylibrary.org/item/18569
page(s): 268; note: name attributed by Fleming to Leach (MS). [details]
original description (of Pupa Draparnaud, 1801) Draparnaud, J. P. R. (1801). <i>Tableau des mollusques terrestres et fluviatiles de la France</i>. Montpellier / Paris (Renaud / Bossange, Masson & Besson). 1-116. , available online at https://www.biodiversitylibrary.org/page/28587765 [details]
original description (of Pupilla (Afripupilla) Pilsbry, 1921) Pilsbry, H. A. (1920-1921). <i>Manual of conchology, structural and systematic, with illustrations of the species</i>. Ser. 2, Pulmonata. Vol. 26: Pupillidae (Vertigininae, Pupillinae). pp i-iv, 1-254, pls 1-24. Philadelphia, published by the Conchological Section, Academy of Natural Sciences. [pp. 1-64, pls 1-8, 23 Dec 1920; pp. 65-128, pls 9-13, 13 May 1921; pp. 129-192, pls 14-18, , figs. 1-4, 4 Aug 1921; pp. 193-254, pls 19-24, figs. 1-4, 21 Dec 1921]. , available online at https://www.biodiversitylibrary.org/page/1295891
page(s): 153 [details]
original description (of Pupilla (Fragipupilla) Schileyko, 1984) Schileyko, A. A. (1984). Nazemnye molljuski podotrjada Pupillina fauny SSSR (Gastropoda, Pulmonata, Geophila). In:. <em>Fauna SSSR, Molljuski.</em> III, 3 [= N.S. 130]: 1-399. Leningrad (Nauka) [14 June]. [Terrestrial molluscs of the suborder Pupillina of the USSR fauna (Gastropoda, Pulmonata, Geophila].
page(s): 191 [details] Available for editors
[request]
original description (of Pupilla (Striopupilla) Pilsbry, 1921) Pilsbry, H. A. (1920-1921). <i>Manual of conchology, structural and systematic, with illustrations of the species</i>. Ser. 2, Pulmonata. Vol. 26: Pupillidae (Vertigininae, Pupillinae). pp i-iv, 1-254, pls 1-24. Philadelphia, published by the Conchological Section, Academy of Natural Sciences. [pp. 1-64, pls 1-8, 23 Dec 1920; pp. 65-128, pls 9-13, 13 May 1921; pp. 129-192, pls 14-18, , figs. 1-4, 4 Aug 1921; pp. 193-254, pls 19-24, figs. 1-4, 21 Dec 1921]. , available online at https://www.biodiversitylibrary.org/page/1295891
page(s): 153, 155 [details]
original description (of Torquatella Held, 1838) Held, F. (1837-1838). Notizen über die Weichthiere Bayerns. Isis (Oken), 30 (4): 303-309 (1837); 30 (12): 901-919 (1838). Leipzig., available online at https://biodiversitylibrary.org/page/27411580
page(s): 919 [details]
original description (of Pupa (Paracoryna) Flach, 1890 †) Flach, K. (1890). Palaeontologische Beiträge. <em>Verhandlungen der Physikalisch-Medicinischen Gesellschaft zu Würzburg, Neue Folge.</em> 24(3): 49-59, pl. 1.
page(s): 52; note: original rank: subsection of Pupa (Coryna) [details]
basis of record Schileyko, A. A. (1998). Treatise on Recent terrestrial pulmonate molluscs. Part 1. Achatinellidae, Amastridae, Orculidae, Strobilopsidae, Spelaeodiscidae, Valloniidae, Cochlicopidae, Pupillidae, Chondrinidae, Pyramidulidae. <em>Ruthenica.</em> Supplement 2: 1-127. Moskva [April].
page(s): 108 [details] Available for editors [request]
additional source Dall, W. H. (1904). Notes on the nomenclature of the Pupacea and associated forms. <em>The Nautilus.</em> 17(10): 114-116., available online at https://www.biodiversitylibrary.org/page/1726537
page(s): 115-116 [details]
additional source Cockerell, T. D. A. (1905). Note on the nomenclature of the snails usually called <i>Pupa</i>. <em>The Nautilus.</em> 18(9): 103-105., available online at https://www.biodiversitylibrary.org/page/1817076 [details]
page(s): 268; note: name attributed by Fleming to Leach (MS). [details]
original description (of Pupa Draparnaud, 1801) Draparnaud, J. P. R. (1801). <i>Tableau des mollusques terrestres et fluviatiles de la France</i>. Montpellier / Paris (Renaud / Bossange, Masson & Besson). 1-116. , available online at https://www.biodiversitylibrary.org/page/28587765 [details]
original description (of Pupilla (Afripupilla) Pilsbry, 1921) Pilsbry, H. A. (1920-1921). <i>Manual of conchology, structural and systematic, with illustrations of the species</i>. Ser. 2, Pulmonata. Vol. 26: Pupillidae (Vertigininae, Pupillinae). pp i-iv, 1-254, pls 1-24. Philadelphia, published by the Conchological Section, Academy of Natural Sciences. [pp. 1-64, pls 1-8, 23 Dec 1920; pp. 65-128, pls 9-13, 13 May 1921; pp. 129-192, pls 14-18, , figs. 1-4, 4 Aug 1921; pp. 193-254, pls 19-24, figs. 1-4, 21 Dec 1921]. , available online at https://www.biodiversitylibrary.org/page/1295891
page(s): 153 [details]
original description (of Pupilla (Fragipupilla) Schileyko, 1984) Schileyko, A. A. (1984). Nazemnye molljuski podotrjada Pupillina fauny SSSR (Gastropoda, Pulmonata, Geophila). In:. <em>Fauna SSSR, Molljuski.</em> III, 3 [= N.S. 130]: 1-399. Leningrad (Nauka) [14 June]. [Terrestrial molluscs of the suborder Pupillina of the USSR fauna (Gastropoda, Pulmonata, Geophila].
page(s): 191 [details] Available for editors
[request]original description (of Pupilla (Striopupilla) Pilsbry, 1921) Pilsbry, H. A. (1920-1921). <i>Manual of conchology, structural and systematic, with illustrations of the species</i>. Ser. 2, Pulmonata. Vol. 26: Pupillidae (Vertigininae, Pupillinae). pp i-iv, 1-254, pls 1-24. Philadelphia, published by the Conchological Section, Academy of Natural Sciences. [pp. 1-64, pls 1-8, 23 Dec 1920; pp. 65-128, pls 9-13, 13 May 1921; pp. 129-192, pls 14-18, , figs. 1-4, 4 Aug 1921; pp. 193-254, pls 19-24, figs. 1-4, 21 Dec 1921]. , available online at https://www.biodiversitylibrary.org/page/1295891
page(s): 153, 155 [details]
original description (of Torquatella Held, 1838) Held, F. (1837-1838). Notizen über die Weichthiere Bayerns. Isis (Oken), 30 (4): 303-309 (1837); 30 (12): 901-919 (1838). Leipzig., available online at https://biodiversitylibrary.org/page/27411580
page(s): 919 [details]
original description (of Pupa (Paracoryna) Flach, 1890 †) Flach, K. (1890). Palaeontologische Beiträge. <em>Verhandlungen der Physikalisch-Medicinischen Gesellschaft zu Würzburg, Neue Folge.</em> 24(3): 49-59, pl. 1.
page(s): 52; note: original rank: subsection of Pupa (Coryna) [details]
basis of record Schileyko, A. A. (1998). Treatise on Recent terrestrial pulmonate molluscs. Part 1. Achatinellidae, Amastridae, Orculidae, Strobilopsidae, Spelaeodiscidae, Valloniidae, Cochlicopidae, Pupillidae, Chondrinidae, Pyramidulidae. <em>Ruthenica.</em> Supplement 2: 1-127. Moskva [April].
page(s): 108 [details] Available for editors
[request]additional source Dall, W. H. (1904). Notes on the nomenclature of the Pupacea and associated forms. <em>The Nautilus.</em> 17(10): 114-116., available online at https://www.biodiversitylibrary.org/page/1726537
page(s): 115-116 [details]
additional source Cockerell, T. D. A. (1905). Note on the nomenclature of the snails usually called <i>Pupa</i>. <em>The Nautilus.</em> 18(9): 103-105., available online at https://www.biodiversitylibrary.org/page/1817076 [details]
Present Inaccurate Introduced: alien Containing type locality
