original description
Seninski, K. (1905). Novyya Dannyya o neogenovych' plastach' Yugozapadnogo Zakavkaz'ya. <em>Trudy Obshchestva Estestvoispytateley pri Imperatorskom Yuryevskom Universitete.</em> 16, 1-80.
page(s): 44-45, pl. 1, figs 4-6 [details]
From editor or global species database
Type locality Mokvi layers at river Duab, Georgia; Kimmerian, early Pliocene [details]